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<article language="en">
	<journal>
		<journal_title>Climate of the Past Discussions</journal_title>
		<journal_url>www.clim-past-discuss.net</journal_url>
		<issn>1814-9340</issn>
		<eissn>1814-9359</eissn>
		<volume_number>5</volume_number>
		<issue_number>1</issue_number>
		<publication_year>2009</publication_year>
	</journal>
	<doi>10.5194/cpd-5-709-2009</doi>
	<article_url>http://www.clim-past-discuss.net/5/709/2009/</article_url>
	<abstract_html>http://www.clim-past-discuss.net/5/709/2009/cpd-5-709-2009.html</abstract_html>
	<fulltext_pdf>http://www.clim-past-discuss.net/5/709/2009/cpd-5-709-2009.pdf</fulltext_pdf>
	<start_page>709</start_page>
	<end_page>734</end_page>
	<publication_date>2009-02-27</publication_date>
	<article_title content_type="html">Mid-Tertiary palaeoenvironments in Thailand: pollen evidences</article_title>
	<authors>
		<author numeration="1" affiliations="1,2">
			<name>P. Sepulchre</name>
			<email>pierre.sepulchre@lsce.ipsl.fr</email>
		</author>
		<author numeration="2" affiliations="2">
			<name>D. Jolly</name>
		</author>
		<author numeration="3" affiliations="3">
			<name>S. Ducrocq</name>
		</author>
		<author numeration="4" affiliations="4">
			<name>Y. Chaimanee</name>
		</author>
		<author numeration="5" affiliations="3">
			<name>J.-J. Jaeger</name>
		</author>
	</authors>
	<affiliations>
		<affiliation numeration="1" content_type="html">Laboratoire des Sciences du Climat et de l&apos;Environnement/Institut Pierre Simon Laplace, UMR CEA/CNRS/UVSQ, Orme des Merisiers, bât. 709, 91191 Gif-sur-Yvette cedex, France</affiliation>
		<affiliation numeration="2" content_type="html">Laboratory of Palaeoenvironments, ISE-M, UMR 5554, Case 61, Université Montpellier II, 34095 Montpellier Cedex 05, France</affiliation>
		<affiliation numeration="3" content_type="html">Institut International de Paléoprimatologie, Paléontologie Humaine: Evolution et Paléoenvironnements, UMR 6046 CNRS, Université de Poitiers, 40 avenue du Recteur Pineau, 86022 Poitiers, France</affiliation>
		<affiliation numeration="4" content_type="html">Paleontology Division, Bureau of Paleontology and Museum, Department of Mineral Resources, Rama VI Road, Bangkok 10400, Thailand</affiliation>
	</affiliations>
	<abstract content_type="html">Only few data from South-East Asia document paleoenvironments evolution
during the Cenozoic. Here we analyse palynological records from four sites of
Thailand. The unique site dated from the Oligocene shows a temperate signal,
while younger Miocene records show a tropical signal with variations in the
hydrology of the deposition basin, as well as a temperate extra-local signal
that could be related to a mid-altitude flora. The mid-Miocene Chiang Muan
basin has delivered several occurrences of an opened biotope (&lt;i&gt;Poaceae&lt;/i&gt;)
alternatively replaced by what could be &lt;i&gt;Syzygium&lt;/i&gt; gallery-forests.
Khorat (&amp;minus;9 to &amp;minus;6.5 Ma) pollen records deliver similar results, but
linked to a strong local signal. Regional and local signals remain difficult
to separate, notably because of badly known palynomorphs, but our high
resolution record confirm oceanic data that suggest high variability in
Miocene climate, as well as modelling studies inferring an early onset of the
Asian monsoon.</abstract>
	<references>
		<reference numeration="1" content_type="text"> Benammi, M., Chaimanee, Y., Urrutia-Fucugauchi, J., et al.: Magnetostratigraphic Study of the Continental Sedimentary Sequence of the Chiang Muan Basin, Northern Thailand: Implications for the Age of the First Miocene Hominoids from Thailand, Int. Geol. Rev., 46(7), 646–654, 2004. </reference>
		<reference numeration="2" content_type="text"> Bennami, M., Urrutia-Fucugauchi, J., Alva-Valdivia, M., et al.: Magnetostratigraphy of the Middle Miocene continental sedimentary sequences of the Mae Moh Basin in northern Thailand: evidence for counterclockwise block rotation, Earth Planet. Sci. Lett., 204, 373–383, 2002. </reference>
		<reference numeration="3" content_type="text"> Blasco, F.: Montagnes du sud de l&apos;Inde: Forêts, savanes, écologie, Madras: BNKP 436~pp., Illustrations, maps: 349–389, 1971. </reference>
		<reference numeration="4" content_type="text"> Bonnefille, R. and Riollet, G.: Pollens des savanes d&apos;Afrique orientale, Paris, Editions du CNRS, 256~pp., 1980. </reference>
		<reference numeration="5" content_type="text"> Briais, A., Patriat, P., Tapponnier, P., et al.: Updated interpretation of magnetic anomalies and seafloor spreading stages in the South China Sea-Implications for the Tertiary tectonics of Southeast Asia, J. Geophys. Res., 98(B4), 6299–6328, 1993. </reference>
		<reference numeration="6" content_type="text"> Chaimanee, Y., Jolly, D., Bennami, M., et al.: A Middle Miocene hominoid from Thailand and orangutan origins, Nature, 422, 61–65, 2003. </reference>
		<reference numeration="7" content_type="text"> Chaimanee, Y., Suteethorn, V., Jintasakul, P., et al.: A new orang-utan relative from the Late Miocene of Thailand, Nature, 427(6973), 439–441, 2004. </reference>
		<reference numeration="8" content_type="text"> Chaimanee, Y., Yamee, C., Tian, P., et al.: \textitKhoratpithecus piriyai, a Late Miocene hominoid of Thailand, Am. J. Phys. Anthropol., 131, 311–323, 2006. </reference>
		<reference numeration="9" content_type="text"> Corsiri, R. and Crouch, A.: Mae Moh coal deposit: Electricity Generating Authority of Thailand, Report of Thailand Australia Lignite Mine Development Project, 1, 448, 1985. </reference>
		<reference numeration="10" content_type="text"> Ducrocq, S., Chaimanee, Y., Suteethorn, V., et al.: Ages and paleoenvironment of Miocene mammalian faunas from Thailand, Palaeogeogr. Palaeoecl., 108(1), 149–163, 1994. </reference>
		<reference numeration="11" content_type="text"> Faegri, K. and Iversen, J.: Textbook of pollen analysis, edited by: Faegri, K., Kaland, P. E., and Krzywinski, K., 328~pp., 1989. </reference>
		<reference numeration="12" content_type="text"> Fluteau, F., Ramstein, G., Besse, J., et al.: Simulating the evolution of the Asian and African monsoons during the past 30 Myr using an atmospheric general circulation model, J. Geophys. Res.-Atmos., 104(D10), 11 995–12 018, 1999. </reference>
		<reference numeration="13" content_type="text"> Germeraad, J. H., Hopping, A., Muller, J., et al.: Palynology of tertiary sediments from tropical areas, Rev. Palaebot. Palyno., 6, 189–348, 1968. </reference>
		<reference numeration="14" content_type="text"> Ginsburg, L.: Les faunes Tertiaires du Nord de la Tha\&quot;&amp;#x0131;lande, Mém. Soc. géol. France, 147, 67–69, 1984. </reference>
		<reference numeration="15" content_type="text"> Ginsburg, L., Mein, P., Thomas, H., et al.: The Miocene of Thailand: Recent Contributions to Vertebrate Palaeontology and Stratygraphy, The palaoenvironment of East Asia from Mid-Tertiary, Centre of Asian Studies, University of Hong Kong, 897–907, 1988. </reference>
		<reference numeration="16" content_type="text"> Hay, W. W., Soeding, E., DeConto, R. M., et al.: The Late Cenozoic uplift – climate change paradox, Int. J. Earth Sci., 91(5); 746–774, 2002. </reference>
		<reference numeration="17" content_type="text"> Hooker, J. D.: The Flora of British India, 740~pp., 1875. </reference>
		<reference numeration="18" content_type="text"> Hoorn, C., Ohja, T., Quade, J., et al.: Palynological evidence for vegetation development and climatic change in the Sub-Himalayan Zone (Neogene, Central Nepal), Palaeogeogr. Palaeoecl., 163, 133–161, 2000. </reference>
		<reference numeration="19" content_type="text"> Huang, T.-C.: Pollen Flora of Taiwan, Ching-Hwa Press co., Ltd., 297~pp., 1972. </reference>
		<reference numeration="20" content_type="text"> Jensen, M.: Trees and Fruits of Southeast Asia, Bangkok, Orchid Press, 234~pp., 2001. </reference>
		<reference numeration="21" content_type="text"> Kanzaki, M., Hara, M., Yamakura, T., et al.: Doi Inthanon Forest Dynamics Plot, Thailand, Tropical Forest Diversity and Dynamism: Findings from a Large-Scale Network Forest Dynamics Plots, edited by: Losos, E. C. and Leigh, E. G., The University of Chicago Press, Chicago, Illinois, USA, 474–481, 2004. </reference>
		<reference numeration="22" content_type="text"> Kuchler, A. W. and Sawyer, J. O.: A study of the vegetation near Chiang Mai, Thailand, Trans. Kansas Acad. Sci., 70, 281–347, 1967. </reference>
		<reference numeration="23" content_type="text"> Letouzey, R.: Étude phytogéographique du Cameroun, 511~pp., 1968. </reference>
		<reference numeration="24" content_type="text"> Liu, X. and Yin, Z. Y.: Sensitivity of East Asian monsoon climate to the uplift of the Tibetan Plateau, Palaeogeogr. Palaeoecl., 183(223), 245~pp., 2002. </reference>
		<reference numeration="25" content_type="text"> Maley, J. and Brenac. P.: Vegetation dynamics, palaeoenvironments and climatic changes in the forests of western Cameroon during the last 28 000 years BP, Rev. Palaeobot. Palyno., 99(2), 157–187, 1998. </reference>
		<reference numeration="26" content_type="text"> Marivaux, L., Bocat, L., Chaimanee, Y., et al.: Cynocephalid dermopterans from the Palaeogene of South Asia (Thailand, Myanmar and Pakistan): systematic, evolutionary and palaeobiogeographic implications, Zool. Scr., 35(4), 395–420, 2006. </reference>
		<reference numeration="27" content_type="text"> Marivaux, L., Chaimanee, Y., Yamee, C., et al.: Discovery of \textitFallomus ladakhensis Nanda and Sahni, 1998 (Mammalia, Rodentia, Diatomyidae) in the lignites of Nong Ya Plong (Phetchaburi Province, Thailand): systematic, biochronological and paleoenvironmental implications, Geodiversitas, 26(3), 493–507, 2004. </reference>
		<reference numeration="28" content_type="text"> Maxwell, J. F. and Elliott, S.: Vegetation and Vascular Flora of Doi Sutep-Pui National Park, Northern Thailand, Thai Studies in Biodiversity, 5, 45–46, 2001. </reference>
		<reference numeration="29" content_type="text"> Morley, C. K., Woganan, N., and Sankumarn, N.: Late Oligocene-Recent stress evolution in rift basins of northern and central Thailand : Implications for escape tectonics, Tectonophysics, 334, 115–150, 2001. </reference>
		<reference numeration="30" content_type="text"> Morley, R. J.: Origin and Evolution of Tropical Rain Forests, Chichester, England, John Wiley and Sons Ltd., 362~pp., 2000. </reference>
		<reference numeration="31" content_type="text"> Muller, J.: Palynological evidence on early differenciationof Angiosperms, Biol. Rev., 45(3), 417–450, 1970. </reference>
		<reference numeration="32" content_type="text"> New, M., Lister, D., Hulme, M., et al.: A high-resolution data set of surface climate over global land areas, Climate Res., 21(1), 1–25, 2002. </reference>
		<reference numeration="33" content_type="text"> Ogawa, H., Yoda, K., and Kira, T.: A preliminary survey on the vegetation of Thailand, Nature and Life in Southeast Asia, 1, 21–157, 1961. </reference>
		<reference numeration="34" content_type="text"> Parnell, J. A. N., Simpson, D. A., Moat, J., et al.: Plant collecting spread and densities: their potential impact on biogeographical studies in Thailand, J. Biogeogr., 30(2), 193–209, 2003. </reference>
		<reference numeration="35" content_type="text"> Peigné, S., Chaimanee, Y., Yamee, C., et al.: A new amphicyonid (Mammalia, Carnivora, Amphicyonidae) from the late middle Miocene of northern Thailand and a review of the amphicyonine record in Asia, J. A. Earth Sci., 26(5), 519–532, 2006. </reference>
		<reference numeration="36" content_type="text"> Ramstein, G., Fluteau, F., Besse, J., et al.: Effect of orogeny, plate motion and land sea distribution on Eurasian climate change over the past 30 million years, Nature, 386(6627), 788–795, 1997. </reference>
		<reference numeration="37" content_type="text"> Sepulchre, P., Ramstein, G., Fluteau, F., et al.: Tectonic uplift and Eastern Africa aridification, Science, 313(5792), 1419–1423, 2006. </reference>
		<reference numeration="38" content_type="text"> Songtham, W., Ratanasthien, B., and Mildenhall, D. C.: New species of algae \textitActinastrum lagerheim and \textitClosterium nitzsch ex Ralfs from Middle Miocene Sediments of Chiang Muan basin, Phayao, Thailand, with tropical pollen composition, Science Asia, 30, 171–181, 2004. </reference>
		<reference numeration="39" content_type="text"> Songtham, W., Ratanasthien, B., Mildenhall, D. C., et al.: Oligocene-Miocene climatic changes in northern Thailand resulting from extrusion tectonics of Southeast Asian landmass, Science Asia, 29, 221–233, 2003. </reference>
		<reference numeration="40" content_type="text"> Sookchaloem, D.: Outline of the forest flora in Northeastern Thailand, Proceedings of the 42nd Kasetsart University Annual Conference, Kasetsart, Thailand, 3–6 February 2004, 40–47, 2004. </reference>
		<reference numeration="41" content_type="text"> Sytsma, K. J., Litt, A., Zjhra, M. L., et al.: Clades, Clocks, and Continents: Historical and Biogeographical Analysis of Myrtaceae, Vochysiaceae, and Relatives in the Southern Hemisphere, Int. J. Plant Sci., 165(S4), 85–105, 2004. </reference>
		<reference numeration="42" content_type="text"> Tassy, P., Anupandhanant, P., Ginsburg, L., et al.: A new \textitStegolophodon (Proboscidea, Mammalia) from the early Miocene of northern Thailand, Geobios(Lyon), 25(4), 511–523, 1992. </reference>
		<reference numeration="43" content_type="text"> Vincens, A., Tiercelin, J. J., Buchet J., et al.: New Oligocene-early Miocene microflora from the southwestern Turkana Basin Palaeoenvironmental implications in the northern Kenya Rift, Palaeogeogr. Palaeoecl., 239(3–4), 470–486, 2006. </reference>
		<reference numeration="44" content_type="text"> Watanasak, M.: Palynological Zonation of Mid-Tertiary Intermontane Basins in Northern Thailand, Chiang Mai University, 216–225, 1989. </reference>
		<reference numeration="45" content_type="text"> Watson, L. and Dallwitz, M. J.: &quot;The families of flowering plants: descriptions, illustrations, identification, and information retrieval&quot;, Retrieved 14 December 2006, \mboxavailable at: http://delta-intkey.com, 2006 </reference>
		<reference numeration="46" content_type="text"> Westerhold, T., Bickert, T., Röhl, U., et al.: Middle to late Miocene oxygen isotope stratigraphy of ODP site 1085 (SE Atlantic): new constrains on Miocene climate variability and sea-level fluctuations, Palaeogeogr. Palaeoecl., 217(3–4), 205–222, 2005. </reference>
		<reference numeration="47" content_type="text"> Wilson, P. G., O&apos;Brien, M. M., Heslewood, M. M., et al.: Relationships within Myrtaceae sensu lato based on a matK phylogeny, Plant Syst. Evol., 251(1), 3–19, 2005. </reference>
		<reference numeration="48" content_type="text"> Zachos, J., Pagani, M., Sloan, L. C., et al.: Trends, Rhythms, and Aberrations in Global Climate 65 Ma to Present, Science, 292, 686–693, 2001. </reference>
	</references>
</article>

